How to compare statistical significance between four subsets of dataframe

General
#1

Comparing roots, shoots and calluses for statistical significance between all four medium types, have subsetted by medium

dat<-read.csv("~/RECAP/Book5.csv",header=T, stringsAsFactors = T)
dat$Medium<-as.factor(dat$Medium) # makes medium a catergorical dataset
M1<-subset(dat,dat$Medium=="M1") # subset into medium types
M2<- subset(dat,dat$Medium=='M2')
M3<- subset(dat,dat$Medium=='M3')
M4<- subset(dat,dat$Medium=='M4')

#2

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